Protein: IPO9
Importin-9 (Ran-binding protein 9)

Gene: IPO9
Organism: Drosophila melanogaster
Interactions: 163
UniProt ID: Q9VGP4

Gene Ontology (top 5):
GO:0005654 - nucleoplasm
GO:0005829 - cytosol
GO:0006606 - protein import into nucleus
GO:0031144 - proteasome localization
GO:0031267 - small GTPase binding

Download interactions: TXT   MITAB-2.5

Interaction Interactor Experiments Homologs Category Interaction
Score
Confidence

782776

2AAA
Serine/threonine-protein phosphatase PP2A 65 kDa regulatory subunit

2

7

High-throughput

0.737

High

1457444

XPO1
Exportin-1

2

12

High-throughput

0.737

High

1465452

Q9V455
Importin subunit alpha

2

20

High-throughput

0.737

High

1474979

Q9VN44
FI07923p

2

17

High-throughput

0.737

High

1463763

RAB5
Ras-related protein Rab5

1

2

High-throughput

0.618

High

1474980

RHEB
GTP-binding protein Rheb homolog

1

2

High-throughput

0.618

High

1267720

Q7KMQ0
26S proteasome regulatory subunit 7

2

5

High-throughput

0.571

High

1474973

NFS1
Cysteine desulfurase, mitochondrial

2

2

High-throughput

0.571

High

1474995

ACADM
Medium-chain specific acyl-CoA dehydrogenase, mitochondrial

2

1

High-throughput

0.571

High

1474996

Q9VSD6
D-Importin 7/RanBP7

2

13

High-throughput

0.571

High

1474966

Q9VI53
LD44267p

2

16

High-throughput

0.549

High

1475004

Q9VW54
26S proteasome non-ATPase regulatory subunit 2

2

0

High-throughput

0.547

High

565029

ATNA
Sodium/potassium-transporting ATPase subunit alpha

2

3

High-throughput

0.521

High

652093

ATC1
Calcium-transporting ATPase sarcoplasmic/endoplasmic reticulum type

2

3

High-throughput

0.521

High

704391

SYEP
Bifunctional glutamate/proline--tRNA ligase

2

3

High-throughput

0.521

High

715150

CLH
Clathrin heavy chain

2

1

High-throughput

0.521

High

852222

COPB
Coatomer subunit beta

2

12

High-throughput

0.521

High

1069190

Q0E8E8
Phosphate carrier protein, mitochondrial

2

1

High-throughput

0.521

High

1265831

Q7K3J0
T-complex protein 1 subunit theta

2

1

High-throughput

0.521

High

1268181

Q7KN94
Electron transfer flavoprotein subunit alpha

2

1

High-throughput

0.521

High

1346268

Q8T8W3
AT21416p

2

3

High-throughput

0.521

High

1465611

Q9V470
Succinate--CoA ligase [GDP-forming] subunit beta, mitochondrial

2

2

High-throughput

0.521

High

1468472

Q9VAW3
glutamine--fructose-6-phosphate transaminase

2

1

High-throughput

0.521

High

1468638

Q9VAY2
Heat shock protein 83

2

11

High-throughput

0.521

High

1474975

CAND1
Cullin-associated NEDD8-dissociated protein 1

2

11

High-throughput

0.521

High

1474987

Q9VQL7
Fatty acid synthase

2

1

High-throughput

0.521

High

1474994

Q9VS57
RE14391p

2

2

High-throughput

0.521

High

1475006

IDH3A
Probable isocitrate dehydrogenase [NAD] subunit alpha, mitochondrial

2

4

High-throughput

0.521

High

1475009

Q9VWL0
Mec2

2

11

High-throughput

0.521

High

1475012

Q9VXQ5
Chaperonin containing TCP1 subunit 6

2

1

High-throughput

0.521

High

304144

RAB7
Ras-related protein Rab7

1

2

High-throughput

0.514

High

355976

O96692
small monomeric GTPase

1

2

High-throughput

0.514

High

1344438

Q8SYG3
E2 ubiquitin-conjugating enzyme

1

6

High-throughput

0.514

High

1440380

Q9I7L0
LP01847p

1

16

High-throughput

0.489

High

1474962

Q9VGU7
Arrestin C-terminal-like domain-containing protein

1

16

High-throughput

0.489

High

1474965

Q9VI00
FI24305p1

1

16

High-throughput

0.489

High

541229

KRAF1
Raf homolog serine/threonine-protein kinase Raf

1

12

High-throughput

0.479

High

1019250

P91634
phosphatidylinositol 3-kinase

1

3

High-throughput

0.479

High

1172663

SIMA
Protein similar

1

7

High-throughput

0.479

High

1303725

AKT1
RAC serine/threonine-protein kinase

1

7

High-throughput

0.479

High

1375021

Q94533
Ribosomal protein S6 kinase

1

10

High-throughput

0.479

High

1378561

FOXO
Forkhead box protein O

1

11

High-throughput

0.479

High

1474984

GOLP3
Golgi phosphoprotein 3 homolog sauron

1

1

High-throughput

0.479

High

1474990

Q9VR61
non-specific serine/threonine protein kinase

1

12

High-throughput

0.479

High

1474991

Q9VRV1
Fiery mountain, isoform A

1

6

High-throughput

0.479

High

1475020

PDPK1
3-phosphoinositide-dependent protein kinase 1

1

10

High-throughput

0.479

High

51915

PO210
Nuclear pore membrane glycoprotein 210

1

2

High-throughput

0.445

High

55816

A1Z9K0
Uncharacterized protein, isoform B

1

4

High-throughput

0.445

High

58107

A1ZBE9
Methionine--tRNA ligase, cytoplasmic

1

3

High-throughput

0.445

High

438004

PYR1
Multifunctional protein r

1

5

High-throughput

0.445

High

875014

PRS4
26S proteasome regulatory subunit 4

1

6

High-throughput

0.445

High

1234119

SARM1
NAD(+) hydrolase sarm1

1

13

High-throughput

0.445

High

1376859

LDH
L-lactate dehydrogenase

1

1

High-throughput

0.445

High

1464442

Q9V3V6
26S proteasome regulatory complex subunit p50

1

10

High-throughput

0.445

High

1464792

Q9V3Z4
GH11341p

1

1

High-throughput

0.445

High

1464860

Q9V405
26S proteasome regulatory subunit 6B

1

4

High-throughput

0.445

High

1466749

Q9V9Q4
LD43819p

1

1

High-throughput

0.445

High

1474971

Q9VK69
T-complex protein 1 subunit delta

1

1

High-throughput

0.445

High

1474976

Q9VKZ8
Ubiquitin carboxyl-terminal hydrolase

1

5

High-throughput

0.445

High

1474986

Q9VQL1
serine--tRNA ligase

1

5

High-throughput

0.445

High

1474999

FRL
Formin-like protein

1

1

High-throughput

0.445

High

1475001

AIMP2
Probable aminoacyl tRNA synthase complex-interacting multifunctional protein 2

1

5

High-throughput

0.445

High

1475011

SYRC
Probable arginine--tRNA ligase, cytoplasmic

1

2

High-throughput

0.445

High

1475027

SYQ
Probable glutamine--tRNA ligase

1

2

High-throughput

0.445

High

52696

A1Z729
NADP-retinol dehydrogenase

1

3

High-throughput

0.437

High

120358

M9PD17
Gigas, isoform B

1

3

High-throughput

0.437

High

654339

DDX6
ATP-dependent RNA helicase me31b

1

15

High-throughput

0.437

High

1011989

CCD1P
Probable cytochrome P450 12d1 proximal, mitochondrial

1

1

High-throughput

0.437

High

1260766

Q76NQ0
Dolichyl-diphosphooligosaccharide--protein glycosyltransferase subunit 1

1

1

High-throughput

0.437

High

1269251

Q7KTD0
ABC-type glutathione-S-conjugate transporter

1

4

High-throughput

0.437

High

1346516

Q8T9L5
Serine/threonine-protein kinase STK11

1

7

High-throughput

0.437

High

1470219

Q9VCC9
LD23779p

1

7

High-throughput

0.437

High

1475008

Q9VWJ6
FI03615p

1

4

High-throughput

0.437

High

1473030

Q9VEP0
Regulatory protein zeste

1

0

High-throughput

0.426

High

1474169

ELP6
Elongator complex protein 6

1

0

High-throughput

0.426

High

1474963

Q9VHL2
T-complex protein 1 subunit eta

1

0

High-throughput

0.426

High

1474982

NUBP2
Cytosolic Fe-S cluster assembly factor Nubp2 homolog

1

0

High-throughput

0.426

High

1474985

Q9VQI5
Interferon-related developmental regulator 1

1

0

High-throughput

0.426

High

55470

A1Z9E3
Elongation factor Tu

1

1

High-throughput

0.407

High

263030

TENM
Teneurin-m

1

2

High-throughput

0.407

High

263406

PENG
Protein penguin

1

3

High-throughput

0.407

High

304252

ERD2
ER lumen protein-retaining receptor

1

1

High-throughput

0.407

High

829669

ARF4
ADP ribosylation factor 4

1

3

High-throughput

0.407

High

852966

ACTZ
Actin-related protein 1

1

2

High-throughput

0.407

High

875282

TCPG
T-complex protein 1 subunit gamma

1

1

High-throughput

0.407

High

1069766

Q0E931
GTP-binding protein 1

1

9

High-throughput

0.407

High

1070111

Q0E9E2
Pyruvate carboxylase

1

1

High-throughput

0.407

High

1173950

ISWI
Chromatin-remodeling complex ATPase chain Iswi

1

5

High-throughput

0.407

High

1176015

MCM4
DNA replication licensing factor MCM4

1

7

High-throughput

0.407

High

1176321

VATA2
V-type proton ATPase catalytic subunit A isoform 2

1

1

High-throughput

0.407

High

1236889

Q6NP35
Dolichyl-diphosphooligosaccharide--protein glycosyltransferase subunit 1

1

1

High-throughput

0.407

High

1262531

Q7JWI7
Hermes, isoform A

1

8

High-throughput

0.407

High

1263236

Q7JXU4
Nucleolar GTP-binding protein 2

1

2

High-throughput

0.407

High

1268719

CCD1D
Probable cytochrome P450 12d1 distal, mitochondrial

1

1

High-throughput

0.407

High

1344135

Q8SY39
ATP-dependent RNA helicase DDX54

1

12

High-throughput

0.407

High

1346060

Q8T4G5
AFG3 like matrix AAA peptidase subunit 2, isoform A

1

4

High-throughput

0.407

High

1378484

Q95U38
Succinate--CoA ligase [ADP-forming] subunit beta, mitochondrial

1

2

High-throughput

0.407

High

1378640

Q95WY3
Nucleolar protein 56

1

6

High-throughput

0.407

High

1465180

Q9V434
Asparagine--tRNA ligase, cytoplasmic

1

2

High-throughput

0.407

High

1467954

Q9VAH7
N-terminal kinase-like protein

1

7

High-throughput

0.407

High

1469671

Q9VBU5
3-dehydrosphinganine reductase

1

2

High-throughput

0.407

High

1471242

DDX18
Probable ATP-dependent RNA helicase pitchoune

1

15

High-throughput

0.407

High

1471672

Q9VDK7
Signal recognition particle subunit SRP72

1

2

High-throughput

0.407

High

1472769

Q9VEH0
Aluminum tubes, isoform A

1

12

High-throughput

0.407

High

1474964

ARL8
ADP-ribosylation factor-like protein 8

1

3

High-throughput

0.407

High

1474967

DDX47
ATP-dependent RNA helicase DDX47

1

14

High-throughput

0.407

High

1474970

Q9VK56
ABC-type glutathione-S-conjugate transporter

1

4

High-throughput

0.407

High

1474972

Q9VKC8
FI03495p

1

17

High-throughput

0.407

High

1474974

Q9VKU1
long-chain-fatty-acid--CoA ligase

1

20

High-throughput

0.407

High

1474977

POE
Protein purity of essence

1

2

High-throughput

0.407

High

1474981

MED1
Mediator of RNA polymerase II transcription subunit 1

1

5

High-throughput

0.407

High

1474997

SRP68
Signal recognition particle subunit SRP68

1

2

High-throughput

0.407

High

1475002

Q9VV51
1-Acylglycerol-3-phosphate O-acyltransferase 3, isoform A

1

4

High-throughput

0.407

High

1475007

Q9VWI2
N(Alpha)-acetyltransferase 15/16, isoform A

1

8

High-throughput

0.407

High

1475010

Q9VXG0
Cell division cycle 50, isoform A

1

2

High-throughput

0.407

High

1475013

Q9VXR5
ATP-binding cassette sub-family F member 2

1

4

High-throughput

0.407

High

1475014

RENT1
Regulator of nonsense transcripts 1 homolog

1

4

High-throughput

0.407

High

1475015

Q9VZ88
AT07585p

1

8

High-throughput

0.407

High

1475017

Q9VZU8
Alpha-1,3/1,6-mannosyltransferase ALG2

1

1

High-throughput

0.407

High

1475021

Q9W1H8
acetyl-CoA C-acyltransferase

1

2

High-throughput

0.407

High

1475023

Q9W414
Regulatory particle triple-A ATPase 4

1

5

High-throughput

0.407

High

1475024

MCM7
DNA replication licensing factor Mcm7

1

5

High-throughput

0.407

High

360572

COX2
Cytochrome c oxidase subunit 2

2

0

High-throughput

0.298

High

1173573

OST48
Dolichyl-diphosphooligosaccharide--protein glycosyltransferase 48 kDa subunit

2

0

High-throughput

0.298

High

1237026

Q6NP72
GEO09659p1

2

0

High-throughput

0.298

High

1264537

Q7K110
Dolichyl-diphosphooligosaccharide--protein glycosyltransferase subunit 2

2

0

High-throughput

0.298

High

1265049

JAGN
Protein jagunal

2

0

High-throughput

0.298

High

1343150

Q8STG9
DSec61alpha

2

0

High-throughput

0.298

High

1465373

KRH2
Krueppel homolog 2

2

0

High-throughput

0.298

High

1474968

Q9VJD1
Glucosidase 2 subunit beta

2

0

High-throughput

0.298

High

1474983

Q9VPR1
GH02075p

2

0

High-throughput

0.298

High

1475003

Q9VV75
AT02348p

2

0

High-throughput

0.298

High

1475016

Q9VZL3
very-long-chain enoyl-CoA reductase

2

0

High-throughput

0.298

High

1475022

Q9W3N1
RH59310p

2

0

High-throughput

0.298

High

1265116

C1GLT
Glycoprotein-N-acetylgalactosamine 3-beta-galactosyltransferase 1

1

0

High-throughput

0.265

Low

1345854

Q8T3Z0
Tektin

1

0

High-throughput

0.265

Low

1173209

DS
Protein dachsous

1

0

High-throughput

0.249

Low

1267401

Q7KLX3
Translocon-associated protein subunit delta

1

0

High-throughput

0.249

Low

1474988

OSTC
Putative oligosaccharyltransferase complex subunit CG9662

1

0

High-throughput

0.249

Low

1474992

MELT
Protein melted

1

0

High-throughput

0.249

Low

1475000

Q9VUK7
RE36140p

1

0

Small-scale

0.249

Low

2725

A0A0B4LFU9
Anastral spindle 2, isoform B

1

0

High-throughput

0.232

Low

2786

UEX
Unextended protein

1

0

High-throughput

0.232

Low

53163

A1Z7K8
AaRS-interacting multifunctional protein 1

1

0

High-throughput

0.232

Low

53703

A1Z843
Nodal modulator 3

1

0

High-throughput

0.232

Low

55367

A1Z9E0
diacylglycerol cholinephosphotransferase

1

0

High-throughput

0.232

Low

1071040

Q0KIB3
Histone acetyltransferase type B catalytic subunit

1

0

High-throughput

0.232

Low

1268355

DYM
Dymeclin

1

0

High-throughput

0.232

Low

1321454

Q8MSW0
Isoleucine--tRNA ligase, cytoplasmic

1

0

High-throughput

0.232

Low

1344241

Q8SY53
GH11935p

1

0

High-throughput

0.232

Low

1465126

API5
Apoptosis inhibitor 5 homolog

1

0

High-throughput

0.232

Low

1468065

Q9VAL7
Calnexin 99A, isoform A

1

0

High-throughput

0.232

Low

1470353

Q9VCF8
LD23561p

1

0

High-throughput

0.232

Low

1474969

Q9VK46
Protein YIPF

1

0

High-throughput

0.232

Low

1474978

Q9VM57
Septin interacting protein 2, isoform A

1

0

High-throughput

0.232

Low

1474989

Q9VQX6
LP05054p

1

0

High-throughput

0.232

Low

1474993

Q9VS47
Lysosomal dipeptide transporter MFSD1

1

0

High-throughput

0.232

Low

1474998

ODR4
Protein odr-4 homolog

1

0

High-throughput

0.232

Low

1475005

Q9VW70
FI21813p1

1

0

High-throughput

0.232

Low

1475018

Q9W0E4
Aminopeptidase

1

0

High-throughput

0.232

Low

1475019

SAC1
Phosphatidylinositol-3-phosphatase SAC1

1

0

High-throughput

0.232

Low

1475025

DDB1
DNA damage-binding protein 1

1

0

High-throughput

0.232

Low

1475026

Q9XZ53
dolichyl-diphosphooligosaccharide--protein glycotransferase

1

0

High-throughput

0.232

Low


Service provided by Combinatics, Japan

Last updated: 11Aug2025