Protein: NP1L4
Nucleosome assembly protein 1-like 4 (Nucleosome assembly protein 2) (NAP-2)

Gene: NP1L4, NAP1L4
Organism: Homo sapiens
Interactions: 184
UniProt ID: Q99733

Gene Ontology (top 5):
GO:0000785 - chromatin
GO:0003682 - chromatin binding
GO:0003723 - RNA binding
GO:0005634 - nucleus
GO:0005737 - cytoplasm

Download interactions: TXT   MITAB-2.5

Interaction Interactor Experiments Homologs Category Interaction
Score
Confidence

936842

NP1L1
Nucleosome assembly protein 1-like 1

12

17

High-throughput

0.977

High

419161

H2A1B
Histone H2A type 1-B/E

3

15

Small-scale

0.853

High

1381850

H2B1A
Histone H2B type 1-A

2

19

High-throughput

0.811

High

1163166

H2B2E
Histone H2B type 2-E

2

20

Small-scale

0.799

High

1399192

NP1L5
Nucleosome assembly protein 1-like 5

2

17

High-throughput

0.789

High

502106

H2AZ
Histone H2A.Z

2

19

Small-scale

0.776

High

502496

H2A1
Histone H2A type 1

2

15

Small-scale

0.776

High

1258925

H32
Histone H3.2

2

19

Small-scale

0.768

High

1067642

EP300
Histone acetyltransferase p300

1

10

Small-scale

0.740

High

1406939

MAGD2
Melanoma-associated antigen D2

4

8

Small-scale

0.694

High

1323963

H2B3B
Histone H2B type 3-B

1

20

High-throughput

0.691

High

1406922

UBE2O
(E3-independent) E2 ubiquitin-conjugating enzyme

3

4

Small-scale

0.656

High

1162017

H31T
Histone H3.1t

1

19

Small-scale

0.647

High

1162878

H2A2C
Histone H2A type 2-C

1

15

Small-scale

0.647

High

1406916

NP1L4
Nucleosome assembly protein 1-like 4

1

16

Small-scale

0.647

High

396798

ESR1
Estrogen receptor

3

8

Small-scale

0.641

High

1057867

RL18
Large ribosomal subunit protein eL18

2

3

High-throughput

0.628

High

449776

H2B1J
Histone H2B type 1-J

1

20

High-throughput

0.618

High

754959

H2B1B
Histone H2B type 1-B

1

20

High-throughput

0.618

High

1406938

TSYL4
Testis-specific Y-encoded-like protein 4

1

18

High-throughput

0.618

High

1114114

CUL3
Cullin-3

2

11

High-throughput

0.611

High

831308

CCNF
Cyclin-F

2

4

High-throughput

0.601

High

1316868

ASPM
Abnormal spindle-like microcephaly-associated protein

2

8

Small-scale

0.601

High

980208

H4
Histone H4

2

5

Small-scale

0.595

High

242707

PRKN
E3 ubiquitin-protein ligase parkin

2

3

High-throughput

0.584

High

315934

RECQ4
ATP-dependent DNA helicase Q4

2

5

High-throughput

0.584

High

644010

FBRL
rRNA 2'-O-methyltransferase fibrillarin

2

5

High-throughput

0.584

High

1123786

DCTN1
Dynactin subunit 1

2

4

High-throughput

0.584

High

1364550

BAP1
Ubiquitin carboxyl-terminal hydrolase BAP1

2

4

Small-scale

0.584

High

911950

JAK3
Tyrosine-protein kinase JAK3

2

4

High-throughput

0.574

High

1051298

KPCZ
Protein kinase C zeta type

2

3

High-throughput

0.571

High

1406923

MYLK2
Myosin light chain kinase 2, skeletal/cardiac muscle

2

3

High-throughput

0.571

High

530277

TAU
Microtubule-associated protein tau

2

3

High-throughput

0.552

High

290125

OFD1
Centriole and centriolar satellite protein OFD1

2

0

High-throughput

0.547

High

1406940

ADIP
Afadin- and alpha-actinin-binding protein

2

0

High-throughput

0.547

High

1291348

LCA5
Lebercilin

2

6

High-throughput

0.537

High

997971

CSK21
Casein kinase II subunit alpha

1

19

Small-scale

0.536

High

134132

IPO5
Importin-5

1

1

High-throughput

0.536

High

334889

IPO7
Importin-7

1

4

High-throughput

0.536

High

342304

BRD1
Bromodomain-containing protein 1

1

2

Small-scale

0.536

High

361341

SODC
Superoxide dismutase [Cu-Zn]

1

1

High-throughput

0.536

High

447353

NPM
Nucleophosmin

1

2

High-throughput

0.536

High

455856

H10
Histone H1.0

1

4

High-throughput

0.536

High

587783

DESP
Desmoplakin

1

2

High-throughput

0.536

High

649398

H1T
Histone H1t

1

5

High-throughput

0.536

High

650572

KAPCG
cAMP-dependent protein kinase catalytic subunit gamma

1

2

High-throughput

0.536

High

783168

ARL3
ADP-ribosylation factor-like protein 3

1

1

High-throughput

0.536

High

846440

MATR3
Matrin-3

1

2

High-throughput

0.536

High

872671

SOX2
Transcription factor SOX-2

1

3

High-throughput

0.536

High

978681

RL23A
Large ribosomal subunit protein uL23

1

4

High-throughput

0.536

High

1106228

MYO9B
Unconventional myosin-IXb

1

5

High-throughput

0.536

High

1111431

DGKZ
Diacylglycerol kinase zeta

1

2

High-throughput

0.536

High

1112679

CUL1
Cullin-1

1

11

High-throughput

0.536

High

1127110

RBM39
RNA-binding protein 39

1

2

High-throughput

0.536

High

1160734

DREB
Drebrin

1

3

High-throughput

0.536

High

1243043

PKN3
Serine/threonine-protein kinase N3

1

5

High-throughput

0.536

High

1291157

CAND1
Cullin-associated NEDD8-dissociated protein 1

1

3

High-throughput

0.536

High

1371847

CUL5
Cullin-5

1

11

High-throughput

0.536

High

1406605

HCD2
3-hydroxyacyl-CoA dehydrogenase type-2

1

2

High-throughput

0.536

High

1406919

SPZ1
Spermatogenic leucine zipper protein 1

1

2

High-throughput

0.536

High

1227338

A16L1
Autophagy-related protein 16-1

2

3

High-throughput

0.533

High

1038580

M3K10
Mitogen-activated protein kinase kinase kinase 10

1

0

High-throughput

0.513

High

1325442

SYNPO
Synaptopodin

1

0

High-throughput

0.513

High

185956

OGT1
UDP-N-acetylglucosamine--peptide N-acetylglucosaminyltransferase 110 kDa subunit

1

5

High-throughput

0.501

High

508503

UBC
Polyubiquitin-C [Cleaved into: Ubiquitin]

1

4

Small-scale

0.501

High

1037332

H11
Histone H1.1

1

5

Small-scale

0.501

High

866801

CAZA2
F-actin-capping protein subunit alpha-2

1

2

High-throughput

0.489

High

867374

CAPZB
F-actin-capping protein subunit beta

1

2

High-throughput

0.489

High

1206958

CEP78
Centrosomal protein of 78 kDa

1

2

High-throughput

0.489

High

1406934

RM35
Large ribosomal subunit protein bL35m

1

1

High-throughput

0.489

High

173270

XPO1
Exportin-1

1

5

High-throughput

0.479

High

187166

PPM1G
Protein phosphatase 1G

1

2

High-throughput

0.479

High

213486

CTIF
CBP80/20-dependent translation initiation factor

1

2

High-throughput

0.479

High

223282

PRC1
Protein regulator of cytokinesis 1

1

1

High-throughput

0.479

High

247415

PHLP1
PH domain leucine-rich repeat-containing protein phosphatase 1

1

6

High-throughput

0.479

High

260359

BRD4
Bromodomain-containing protein 4

1

11

High-throughput

0.479

High

277399

TP4A3
Protein tyrosine phosphatase type IVA 3

1

5

High-throughput

0.479

High

320354

PHF14
PHD finger protein 14

1

1

High-throughput

0.479

High

325071

HEXI1
Protein HEXIM1

1

2

High-throughput

0.479

High

331262

KI20A
Kinesin-like protein KIF20A

1

5

High-throughput

0.479

High

412505

NTRK1
High affinity nerve growth factor receptor

1

3

High-throughput

0.479

High

463727

LYN
Tyrosine-protein kinase Lyn

1

1

High-throughput

0.479

High

537777

HSP7C
Heat shock cognate 71 kDa protein

1

4

High-throughput

0.479

High

650925

ROA2
Heterogeneous nuclear ribonucleoproteins A2/B1

1

1

High-throughput

0.479

High

708866

PTN4
Tyrosine-protein phosphatase non-receptor type 4

1

4

High-throughput

0.479

High

731564

STIP1
Stress-induced-phosphoprotein 1

1

5

High-throughput

0.479

High

764419

CTNA1
Catenin alpha-1

1

1

Small-scale

0.479

High

791657

SYUA
Alpha-synuclein

1

2

High-throughput

0.479

High

862407

YAP1
Transcriptional coactivator YAP1

1

10

High-throughput

0.479

High

887166

CEBPA
CCAAT/enhancer-binding protein alpha

1

4

High-throughput

0.479

High

891298

GSK3A
Glycogen synthase kinase-3 alpha

1

9

High-throughput

0.479

High

891763

GSK3B
Glycogen synthase kinase-3 beta

1

9

High-throughput

0.479

High

905070

MECP2
Methyl-CpG-binding protein 2

1

3

High-throughput

0.479

High

920231

PLK1
Serine/threonine-protein kinase PLK1

1

6

High-throughput

0.479

High

984931

RL31
Large ribosomal subunit protein eL31

1

5

High-throughput

0.479

High

1121317

TRI14
Tripartite motif-containing protein 14

1

4

High-throughput

0.479

High

1138550

U5S1
116 kDa U5 small nuclear ribonucleoprotein component

1

2

High-throughput

0.479

High

1140746

KIF14
Kinesin-like protein KIF14

1

1

High-throughput

0.479

High

1146465

KS6A2
Ribosomal protein S6 kinase alpha-2

1

4

High-throughput

0.479

High

1155699

EZH2
Histone-lysine N-methyltransferase EZH2

1

2

High-throughput

0.479

High

1241758

PRP8
Pre-mRNA-processing-splicing factor 8

1

1

High-throughput

0.479

High

1250645

BCOR
BCL-6 corepressor

1

2

High-throughput

0.479

High

1255230

TRI67
Tripartite motif-containing protein 67

1

11

High-throughput

0.479

High

1258377

MOB3C
MOB kinase activator 3C

1

1

High-throughput

0.479

High

1271812

MEPCE
7SK snRNA methylphosphate capping enzyme

1

3

High-throughput

0.479

High

1276701

PARPT
Protein mono-ADP-ribosyltransferase TIPARP

1

2

High-throughput

0.479

High

1280303

DNJB7
DnaJ homolog subfamily B member 7

1

2

High-throughput

0.479

High

1280436

HUWE1
E3 ubiquitin-protein ligase HUWE1

1

6

High-throughput

0.479

High

1289572

NAA40
N-alpha-acetyltransferase 40

1

1

High-throughput

0.479

High

1291566

HOOK3
Protein Hook homolog 3

1

3

High-throughput

0.479

High

1314217

SPB1
pre-rRNA 2'-O-ribose RNA methyltransferase FTSJ3

1

3

High-throughput

0.479

High

1322435

SPICE
Spindle and centriole-associated protein 1

1

1

High-throughput

0.479

High

1324057

RHG24
Rho GTPase-activating protein 24

1

2

High-throughput

0.479

High

1331549

CE120
Centrosomal protein of 120 kDa

1

7

High-throughput

0.479

High

1331665

RM43
Large ribosomal subunit protein mL43

1

1

High-throughput

0.479

High

1350426

BICD2
Protein bicaudal D homolog 2

1

4

High-throughput

0.479

High

1363955

SMG7
Nonsense-mediated mRNA decay factor SMG7

1

2

High-throughput

0.479

High

1368083

HDAC2
Histone deacetylase 2

1

2

High-throughput

0.479

High

1368363

REQU
Zinc finger protein ubi-d4

1

9

High-throughput

0.479

High

1368579

KAT6A
Histone acetyltransferase KAT6A

1

9

High-throughput

0.479

High

1370571

FGF11
Fibroblast growth factor 11

1

1

High-throughput

0.479

High

1380723

RL36L
Ribosomal protein eL42-like

1

6

High-throughput

0.479

High

1397350

CI072
Guanine nucleotide exchange factor C9orf72

1

3

High-throughput

0.479

High

1403324

RUFY1
RUN and FYVE domain-containing protein 1

1

4

High-throughput

0.479

High

1406920

YTHD1
YTH domain-containing family protein 1

1

3

High-throughput

0.479

High

1406924

CHM4B
Charged multivesicular body protein 4b

1

4

High-throughput

0.479

High

1406925

EGLN3
Prolyl hydroxylase EGLN3

1

2

High-throughput

0.479

High

1406926

HM20A
High mobility group protein 20A

1

7

High-throughput

0.479

High

1406930

PIHD1
PIH1 domain-containing protein 1

1

1

High-throughput

0.479

High

1406931

AKIP
Small ribosomal subunit protein mS38

1

1

High-throughput

0.479

High

1406936

ZRAN1
Ubiquitin thioesterase ZRANB1

1

4

High-throughput

0.479

High

1406941

ZN330
Zinc finger protein 330

1

2

High-throughput

0.479

High

1327153

FBXL6
F-box/LRR-repeat protein 6

1

1

High-throughput

0.470

High

869788

MTR1A
Melatonin receptor type 1A

1

2

High-throughput

0.464

High

630007

BTF3
Transcription factor BTF3

1

0

High-throughput

0.458

High

1037886

HEN1
Helix-loop-helix protein 1

1

0

High-throughput

0.458

High

1156831

E2F4
Transcription factor E2F4

1

0

High-throughput

0.458

High

1161799

CDKN3
Cyclin-dependent kinase inhibitor 3

1

0

High-throughput

0.458

High

1214563

ACBD5
Acyl-CoA-binding domain-containing protein 5

1

0

High-throughput

0.458

High

1228317

UTP25
U3 small nucleolar RNA-associated protein 25 homolog

1

0

High-throughput

0.458

High

1334821

SERB1
SERPINE1 mRNA-binding protein 1

1

0

High-throughput

0.458

High

1406928

DUS22
Dual specificity protein phosphatase 22

1

0

High-throughput

0.458

High

1406932

RM39
Large ribosomal subunit protein mL39

1

0

High-throughput

0.458

High

1406933

F120A
Constitutive coactivator of PPAR-gamma-like protein 1

1

0

High-throughput

0.458

High

1406937

TES
Testin

1

0

High-throughput

0.458

High

67031

ZN729
Zinc finger protein 729

1

3

High-throughput

0.445

High

184650

MK13
Mitogen-activated protein kinase 13

1

3

Small-scale

0.445

High

430277

JUN
Transcription factor Jun

1

5

High-throughput

0.445

High

844858

EAA1
Excitatory amino acid transporter 1

1

1

High-throughput

0.445

High

935047

TERA
Transitional endoplasmic reticulum ATPase

1

2

High-throughput

0.445

High

952307

GBRL2
Gamma-aminobutyric acid receptor-associated protein-like 2

1

6

High-throughput

0.445

High

1209560

LRRK2
Leucine-rich repeat serine/threonine-protein kinase 2

1

5

Small-scale

0.445

High

1243520

CSAG1
Chondrosarcoma-associated gene 1 protein

1

2

High-throughput

0.445

High

1406917

ZN577
Zinc finger protein 577

1

2

High-throughput

0.445

High

106539

APRIO
Alternative prion protein

1

1

High-throughput

0.437

High

127720

TR10A
Tumor necrosis factor receptor superfamily member 10A

1

3

High-throughput

0.437

High

328726

VAMP5
Vesicle-associated membrane protein 5

1

1

High-throughput

0.437

High

340223

PCNT
Pericentrin

1

7

High-throughput

0.437

High

341133

DIRA3
GTP-binding protein Di-Ras3

1

3

High-throughput

0.437

High

519207

COX8A
Cytochrome c oxidase subunit 8A, mitochondrial

1

2

High-throughput

0.437

High

945080

RAB25
Ras-related protein Rab-25

1

1

High-throughput

0.437

High

976513

RB11A
Ras-related protein Rab-11A

1

1

High-throughput

0.437

High

1049195

RON
Macrophage-stimulating protein receptor

1

2

High-throughput

0.437

High

1063566

GOGA2
Golgin subfamily A member 2

1

4

High-throughput

0.437

High

1142156

P2RY6
P2Y purinoceptor 6

1

3

High-throughput

0.437

High

1229053

RNF43
E3 ubiquitin-protein ligase RNF43

1

1

High-throughput

0.437

High

1246951

RICTR
Rapamycin-insensitive companion of mTOR

1

3

High-throughput

0.437

High

1385840

CHM4C
Charged multivesicular body protein 4c

1

4

High-throughput

0.437

High

1396182

RPGR1
X-linked retinitis pigmentosa GTPase regulator-interacting protein 1

1

7

High-throughput

0.437

High

1406918

CP250
Centrosome-associated protein CEP250

1

2

High-throughput

0.437

High

1406927

ANLN
Anillin

1

2

High-throughput

0.437

High

1406929

KBTB4
Kelch repeat and BTB domain-containing protein 4

1

1

High-throughput

0.437

High

1406935

PD1L1
Programmed cell death 1 ligand 1

1

1

High-throughput

0.437

High

1326050

RYBP
RING1 and YY1-binding protein

1

0

High-throughput

0.426

High

1406921

ZDHC5
Palmitoyltransferase ZDHHC5

1

0

High-throughput

0.426

High

1288443

SO6A1
Solute carrier organic anion transporter family member 6A1

1

1

High-throughput

0.407

High

1335690

MOT11
Monocarboxylate transporter 11

1

1

High-throughput

0.407

High

1352093

Q8TE30
RNA-directed DNA polymerase

1

2

Small-scale

0.407

High

1387439

CMBL
Carboxymethylenebutenolidase homolog

1

3

High-throughput

0.407

High

1391021

OTUB1
Ubiquitin thioesterase OTUB1

1

1

High-throughput

0.407

High

858200

BKRB1
B1 bradykinin receptor

1

0

High-throughput

0.282

High

1178333

CL16A
Protein CLEC16A

1

0

High-throughput

0.249

Low

1308369

CCD71
Coiled-coil domain-containing protein 71

1

0

Small-scale

0.249

Low


Service provided by Combinatics, Japan

Last updated: 11Aug2025