Protein: IPO9
Importin-9 (Imp9) (Ran-binding protein 9) (RanBP9)

Gene: IPO9
Organism: Homo sapiens
Interactions: 184
UniProt ID: Q96P70

Gene Ontology (top 5):
GO:0005635 - nuclear envelope
GO:0005654 - nucleoplasm
GO:0005737 - cytoplasm
GO:0005829 - cytosol
GO:0006606 - protein import into nucleus

Download interactions: TXT   MITAB-2.5

Interaction Interactor Experiments Homologs Category Interaction
Score
Confidence

981648

RAN
GTP-binding nuclear protein Ran

3

20

Small-scale

0.866

High

1381845

H2B1A
Histone H2B type 1-A

3

14

High-throughput

0.861

High

256958

H2B1K
Histone H2B type 1-K

3

15

High-throughput

0.827

High

981025

H2B1C
Histone H2B type 1-C/E/F/G/I

3

15

High-throughput

0.827

High

962392

RHOA
Transforming protein RhoA

3

14

High-throughput

0.817

High

591795

H2AX
Histone H2AX

2

9

High-throughput

0.811

High

1399504

H2AW
Core histone macro-H2A.2

2

10

High-throughput

0.811

High

1163148

H2B2E
Histone H2B type 2-E

2

15

High-throughput

0.789

High

1323959

H2B3B
Histone H2B type 3-B

2

15

High-throughput

0.789

High

631418

RAB5A
Ras-related protein Rab-5A

2

1

High-throughput

0.737

High

1015689

RHOG
Rho-related GTP-binding protein RhoG

2

12

Small-scale

0.737

High

845471

PE2R3
Prostaglandin E2 receptor EP3 subtype

3

14

High-throughput

0.667

High

1291662

CMTD1
Catechol O-methyltransferase domain-containing protein 1

3

8

High-throughput

0.667

High

1399505

VTI1B
Vesicle transport through interaction with t-SNAREs homolog 1B

3

1

High-throughput

0.667

High

1399511

TN13B
Tumor necrosis factor ligand superfamily member 13B

3

8

High-throughput

0.667

High

1330735

SIGL1
SIGLEC family-like protein 1

3

6

High-throughput

0.662

High

1199886

AKIR2
Akirin-2

4

0

Small-scale

0.661

High

372884

MYC
Myc proto-oncogene protein

3

6

High-throughput

0.656

High

564238

CFTR
Cystic fibrosis transmembrane conductance regulator

3

10

High-throughput

0.650

High

966763

1433G
14-3-3 protein gamma

2

14

High-throughput

0.628

High

1399512

KLDC2
Kelch domain-containing protein 2

2

3

High-throughput

0.628

High

419144

H2A1B
Histone H2A type 1-B/E

1

9

High-throughput

0.618

High

502100

H2AZ
Histone H2A.Z

1

10

High-throughput

0.618

High

502485

H2A1
Histone H2A type 1

1

9

High-throughput

0.618

High

1162871

H2A2C
Histone H2A type 2-C

1

9

High-throughput

0.618

High

1232714

H2A2A
Histone H2A type 2-A

1

9

High-throughput

0.618

High

1399494

H2AJ
Histone H2A.J

1

9

High-throughput

0.618

High

659933

H2B1O
Histone H2B type 1-O

1

15

High-throughput

0.618

High

946859

H2B1D
Histone H2B type 1-D

1

15

High-throughput

0.618

High

955489

CDC42
Cell division control protein 42 homolog

1

13

High-throughput

0.618

High

1208876

H2B2F
Histone H2B type 2-F

1

15

High-throughput

0.618

High

1372277

H2B1H
Histone H2B type 1-H

1

15

High-throughput

0.618

High

1399490

H2B1M
Histone H2B type 1-M

1

15

High-throughput

0.618

High

1399491

H2B1L
Histone H2B type 1-L

1

14

High-throughput

0.618

High

1326214

PNKD
Probable thioesterase PNKD

2

5

High-throughput

0.611

High

1399501

DISC1
Disrupted in schizophrenia 1 protein

2

3

High-throughput

0.606

High

363732

EGFR
Epidermal growth factor receptor

2

11

Small-scale

0.584

High

1123774

DCTN1
Dynactin subunit 1

2

3

High-throughput

0.584

High

466952

ACM4
Muscarinic acetylcholine receptor M4

2

11

High-throughput

0.574

High

630186

ACM3
Muscarinic acetylcholine receptor M3

2

10

High-throughput

0.574

High

1399498

PACC1
Proton-activated chloride channel

2

6

High-throughput

0.574

High

1097155

MK07
Mitogen-activated protein kinase 7

2

5

High-throughput

0.571

High

1227293

A16L1
Autophagy-related protein 16-1

2

6

High-throughput

0.571

High

1370801

BAD
Bcl2-associated agonist of cell death

2

2

High-throughput

0.571

High

220000

XRCC3
DNA repair protein XRCC3

1

2

High-throughput

0.536

High

351947

S1PR4
Sphingosine 1-phosphate receptor 4

1

14

High-throughput

0.536

High

638229

S1PR1
Sphingosine 1-phosphate receptor 1

1

12

High-throughput

0.536

High

690352

STOM
Stomatin

1

9

High-throughput

0.536

High

691345

1433T
14-3-3 protein theta

1

14

High-throughput

0.536

High

730159

1433B
14-3-3 protein beta/alpha

1

14

High-throughput

0.536

High

733508

GP183
G-protein coupled receptor 183

1

20

High-throughput

0.536

High

843889

HD
Huntingtin

1

4

High-throughput

0.536

High

849396

RANG
Ran-specific GTPase-activating protein

1

16

High-throughput

0.536

High

935951

ECHB
Trifunctional enzyme subunit beta, mitochondrial

1

2

High-throughput

0.536

High

971896

1433E
14-3-3 protein epsilon

1

2

High-throughput

0.536

High

990622

1433Z
14-3-3 protein zeta/delta

1

14

High-throughput

0.536

High

1014180

CBX1
Chromobox protein homolog 1

1

1

High-throughput

0.536

High

1022938

EFNB1
Ephrin-B1

1

6

High-throughput

0.536

High

1049681

1433F
14-3-3 protein eta

1

14

High-throughput

0.536

High

1111205

SNW1
SNW domain-containing protein 1

1

2

High-throughput

0.536

High

1124525

MAP7
Ensconsin

1

11

High-throughput

0.536

High

1161189

MK06
Mitogen-activated protein kinase 6

1

8

High-throughput

0.536

High

1238249

CK087
Uncharacterized protein C11orf87

1

1

High-throughput

0.536

High

1257887

UBP49
Ubiquitin carboxyl-terminal hydrolase 49

1

2

High-throughput

0.536

High

1336058

ARRD4
Arrestin domain-containing protein 4

1

12

High-throughput

0.536

High

1384108

ARRD3
Arrestin domain-containing protein 3

1

12

High-throughput

0.536

High

1399487

SEC62
Translocation protein SEC62

1

2

High-throughput

0.536

High

1399488

CDC5L
Cell division cycle 5-like protein

1

6

High-throughput

0.536

High

1399509

FZD10
Frizzled-10

1

7

High-throughput

0.536

High

1218795

BROX
BRO1 domain-containing protein BROX

1

0

High-throughput

0.513

High

1399489

SORT
Sortilin

1

0

High-throughput

0.513

High

422931

A4
Amyloid-beta precursor protein

1

3

High-throughput

0.501

High

166018

FFAR1
Free fatty acid receptor 1

1

20

High-throughput

0.489

High

253885

TPST2
Protein-tyrosine sulfotransferase 2

1

3

High-throughput

0.489

High

702830

5HT2C
5-hydroxytryptamine receptor 2C

1

9

High-throughput

0.489

High

829502

ECHA
Trifunctional enzyme subunit alpha, mitochondrial

1

1

High-throughput

0.489

High

1142148

P2RY6
P2Y purinoceptor 6

1

20

High-throughput

0.489

High

1249833

BTNL9
Butyrophilin-like protein 9

1

6

High-throughput

0.489

High

1356859

PTPM1
Phosphatidylglycerophosphatase and protein-tyrosine phosphatase 1

1

10

High-throughput

0.489

High

1386325

FAF2
FAS-associated factor 2

1

4

High-throughput

0.489

High

1399513

FBX7
F-box only protein 7

1

0

High-throughput

0.480

High

294203

CD123
Translation initiation factor eIF2 assembly protein

1

3

High-throughput

0.479

High

315902

RECQ4
ATP-dependent DNA helicase Q4

1

7

High-throughput

0.479

High

340215

PCNT
Pericentrin

1

5

High-throughput

0.479

High

350114

SNAI1
Zinc finger protein SNAI1

1

20

High-throughput

0.479

High

401322

RAF1
RAF proto-oncogene serine/threonine-protein kinase

1

8

High-throughput

0.479

High

404341

GCR
Glucocorticoid receptor

1

10

High-throughput

0.479

High

437307

K2C8
Keratin, type II cytoskeletal 8

1

3

High-throughput

0.479

High

521414

ARAF
Serine/threonine-protein kinase A-Raf

1

8

High-throughput

0.479

High

535426

BIP
Endoplasmic reticulum chaperone BiP

1

6

High-throughput

0.479

High

537738

HSP7C
Heat shock cognate 71 kDa protein

1

3

High-throughput

0.479

High

589452

RFA2
Replication protein A 32 kDa subunit

1

6

High-throughput

0.479

High

643994

FBRL
rRNA 2'-O-methyltransferase fibrillarin

1

2

High-throughput

0.479

High

658978

PTPRE
Receptor-type tyrosine-protein phosphatase epsilon

1

9

High-throughput

0.479

High

731547

STIP1
Stress-induced-phosphoprotein 1

1

8

High-throughput

0.479

High

768930

RFA3
Replication protein A 14 kDa subunit

1

5

High-throughput

0.479

High

831267

CCNF
Cyclin-F

1

8

High-throughput

0.479

High

879374

NR2C2
Nuclear receptor subfamily 2 group C member 2

1

2

High-throughput

0.479

High

975184

ARF6
ADP-ribosylation factor 6

1

2

High-throughput

0.479

High

1004991

RNF4
E3 ubiquitin-protein ligase RNF4

1

5

High-throughput

0.479

High

1067613

EP300
Histone acetyltransferase p300

1

14

High-throughput

0.479

High

1096620

AIMP2
Aminoacyl tRNA synthase complex-interacting multifunctional protein 2

1

4

High-throughput

0.479

High

1103326

CTBP1
C-terminal-binding protein 1

1

1

High-throughput

0.479

High

1107117

BIRC3
Baculoviral IAP repeat-containing protein 3

1

8

High-throughput

0.479

High

1114050

CUL3
Cullin-3

1

10

High-throughput

0.479

High

1115009

CUL4A
Cullin-4A

1

10

High-throughput

0.479

High

1146458

KS6A2
Ribosomal protein S6 kinase alpha-2

1

8

High-throughput

0.479

High

1152458

T22D1
TSC22 domain family protein 1

1

2

High-throughput

0.479

High

1163399

CAH9
Carbonic anhydrase 9

1

6

Small-scale

0.479

High

1218636

OTU1
Ubiquitin thioesterase OTU1

1

3

High-throughput

0.479

High

1228456

FTM
Protein fantom

1

14

High-throughput

0.479

High

1229633

VIR
Protein virilizer homolog

1

5

High-throughput

0.479

High

1238462

ZN746
Zinc finger protein 746

1

20

High-throughput

0.479

High

1246938

RICTR
Rapamycin-insensitive companion of mTOR

1

3

High-throughput

0.479

High

1255127

TRI67
Tripartite motif-containing protein 67

1

9

High-throughput

0.479

High

1287005

UBP37
Ubiquitin carboxyl-terminal hydrolase 37

1

2

Small-scale

0.479

High

1287415

SYVN1
E3 ubiquitin-protein ligase synoviolin

1

7

Small-scale

0.479

High

1312411

ASXL1
Polycomb group protein ASXL1

1

5

High-throughput

0.479

High

1334794

SERB1
SERPINE1 mRNA-binding protein 1

1

6

High-throughput

0.479

High

1357521

DCAF4
DDB1- and CUL4-associated factor 4

1

3

High-throughput

0.479

High

1363823

GBF1
Golgi-specific brefeldin A-resistance guanine nucleotide exchange factor 1

1

2

High-throughput

0.479

High

1364527

BAP1
Ubiquitin carboxyl-terminal hydrolase BAP1

1

5

High-throughput

0.479

High

1380094

FBXW7
F-box/WD repeat-containing protein 7

1

4

High-throughput

0.479

High

1380628

FBX32
F-box only protein 32

1

2

High-throughput

0.479

High

1393605

DDRGK
DDRGK domain-containing protein 1

1

5

High-throughput

0.479

High

1396180

RPGR1
X-linked retinitis pigmentosa GTPase regulator-interacting protein 1

1

14

High-throughput

0.479

High

1397093

CEP19
Centrosomal protein of 19 kDa

1

1

High-throughput

0.479

High

1399497

SPRTN
DNA-dependent metalloprotease SPRTN

1

1

High-throughput

0.479

High

1399499

ECT2
Protein ECT2

1

7

High-throughput

0.479

High

1399503

TLS1
Splicing factor C9orf78

1

6

High-throughput

0.479

High

1399506

ZRAN1
Ubiquitin thioesterase ZRANB1

1

13

High-throughput

0.479

High

1399510

TRI33
E3 ubiquitin-protein ligase TRIM33

1

10

High-throughput

0.479

High

1399514

ZN330
Zinc finger protein 330

1

3

High-throughput

0.479

High

1399515

AAKB1
5'-AMP-activated protein kinase subunit beta-1

1

1

High-throughput

0.479

High

1399517

NEMO
NF-kappa-B essential modulator

1

3

High-throughput

0.479

High

869783

MTR1A
Melatonin receptor type 1A

1

11

High-throughput

0.464

High

880951

MTR1B
Melatonin receptor type 1B

1

8

High-throughput

0.464

High

447329

NPM
Nucleophosmin

1

0

High-throughput

0.458

High

1204830

DCA17
DDB1- and CUL4-associated factor 17

1

0

High-throughput

0.458

High

1399500

CPAP
Centrosomal P4.1-associated protein

1

0

High-throughput

0.458

High

104210

F5H1C8
Solute carrier family 15 member 3

1

9

High-throughput

0.445

High

171559

S27A2
Long-chain fatty acid transport protein 2

1

20

High-throughput

0.445

High

190249

MOT4
Monocarboxylate transporter 4

1

8

High-throughput

0.445

High

292279

S22A3
Solute carrier family 22 member 3

1

2

High-throughput

0.445

High

430252

JUN
Transcription factor Jun

1

9

High-throughput

0.445

High

530239

TAU
Microtubule-associated protein tau

1

8

High-throughput

0.445

High

854575

S15A1
Solute carrier family 15 member 1

1

9

High-throughput

0.445

High

1022829

RHG04
Rho GTPase-activating protein 4

1

5

High-throughput

0.445

High

1130395

RRP1B
Ribosomal RNA processing protein 1 homolog B

1

2

High-throughput

0.445

High

1209531

LRRK2
Leucine-rich repeat serine/threonine-protein kinase 2

1

6

High-throughput

0.445

High

1240936

S27A4
Long-chain fatty acid transport protein 4

1

20

High-throughput

0.445

High

1243517

CSAG1
Chondrosarcoma-associated gene 1 protein

1

3

High-throughput

0.445

High

1243653

S27A1
Long-chain fatty acid transport protein 1

1

20

High-throughput

0.445

High

1309398

SPNS2
Sphingosine-1-phosphate transporter SPNS2

1

2

High-throughput

0.445

High

1313978

S35F3
Solute carrier family 35 member F3

1

1

High-throughput

0.445

High

1315547

ULK2
Serine/threonine-protein kinase ULK2

1

2

High-throughput

0.445

High

1316863

ASPM
Abnormal spindle-like microcephaly-associated protein

1

2

Small-scale

0.445

High

1325900

MFD4A
Major facilitator superfamily domain-containing protein 4A

1

4

High-throughput

0.445

High

1399496

ZDHC5
Palmitoyltransferase ZDHHC5

1

3

High-throughput

0.445

High

1399507

AGO2
Protein argonaute-2

1

5

High-throughput

0.445

High

1399508

AGO1
Protein argonaute-1

1

5

High-throughput

0.445

High

106492

APRIO
Alternative prion protein

1

6

High-throughput

0.437

High

136964

AGRIN
Agrin [Cleaved into: Agrin N-terminal 110 kDa subunit; Agrin C-terminal 110 kDa subunit; Agrin C-terminal 90 kDa fragment

1

2

High-throughput

0.437

High

412455

NTRK1
High affinity nerve growth factor receptor

1

10

High-throughput

0.437

High

519197

COX8A
Cytochrome c oxidase subunit 8A, mitochondrial

1

2

High-throughput

0.437

High

650368

FGFR3
Fibroblast growth factor receptor 3

1

11

High-throughput

0.437

High

720303

UFO
Tyrosine-protein kinase receptor UFO

1

5

High-throughput

0.437

High

916326

TXTP
Tricarboxylate transport protein, mitochondrial

1

2

High-throughput

0.437

High

928651

PMS1
PMS1 protein homolog 1

1

2

High-throughput

0.437

High

1126154

GRM2
Metabotropic glutamate receptor 2

1

1

High-throughput

0.437

High

1249131

LMBRL
Protein LMBR1L

1

5

High-throughput

0.437

High

1271800

MEPCE
7SK snRNA methylphosphate capping enzyme

1

1

High-throughput

0.437

High

1276660

PARPT
Protein mono-ADP-ribosyltransferase TIPARP

1

5

High-throughput

0.437

High

1313803

S15A3
Solute carrier family 15 member 3

1

9

High-throughput

0.437

High

1399492

ZFY21
Zinc finger FYVE domain-containing protein 21

1

4

Small-scale

0.437

High

1399493

OVOL2
Transcription factor Ovo-like 2

1

20

Small-scale

0.437

High

1399502

B3GN2
N-acetyllactosaminide beta-1,3-N-acetylglucosaminyltransferase 2

1

4

High-throughput

0.437

High

1295388

MF2L2
Probable guanine nucleotide exchange factor MCF2L2

1

0

High-throughput

0.426

High

1106

SIM26
Small integral membrane protein 26

1

6

High-throughput

0.407

High

1288439

SO6A1
Solute carrier organic anion transporter family member 6A1

1

8

High-throughput

0.407

High

1335680

MOT11
Monocarboxylate transporter 11

1

8

High-throughput

0.407

High

1399516

CSAG2
Chondrosarcoma-associated gene 2/3 protein

1

3

High-throughput

0.407

High

1398604

DAA10
Dynein axonemal assembly factor 10

1

0

High-throughput

0.249

Low

1399495

RPAP1
RNA polymerase II-associated protein 1

1

0

High-throughput

0.249

Low


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Last updated: 11Aug2025