Protein: NEUL4
Neuralized-like protein 4

Gene: NEUL4, NEURL4
Organism: Homo sapiens
Interactions: 229
UniProt ID: Q96JN8

Gene Ontology (top 5):
GO:0005739 - mitochondrion
GO:0005814 - centriole
GO:0007283 - spermatogenesis
GO:0043504 - mitochondrial DNA repair
GO:0061630 - ubiquitin protein ligase activity

Download interactions: TXT   MITAB-2.5

Interaction Interactor Experiments Homologs Category Interaction
Score
Confidence

343015

HERC2
E3 ubiquitin-protein ligase HERC2

8

1

Small-scale

0.744

High

213230

CP110
Centriolar coiled-coil protein of 110 kDa

4

0

Small-scale

0.670

High

177040

CE290
Centrosomal protein of 290 kDa

4

6

Small-scale

0.666

High

1309477

CEP97
Centrosomal protein of 97 kDa

4

0

Small-scale

0.638

High

1092627

ECH1
Delta(3,5)-Delta(2,4)-dienoyl-CoA isomerase, mitochondrial

2

1

High-throughput

0.628

High

1395325

CC136
Coiled-coil domain-containing protein 136

2

3

High-throughput

0.628

High

1050507

UBE3A
Ubiquitin-protein ligase E3A

3

0

Small-scale

0.628

High

1162495

CLPP
ATP-dependent Clp protease proteolytic subunit, mitochondrial

3

0

High-throughput

0.622

High

284603

ECI2
Enoyl-CoA delta isomerase 2

2

1

Small-scale

0.611

High

778756

PRS7
26S proteasome regulatory subunit 7

3

2

High-throughput

0.609

High

127166

APBB1
Amyloid beta precursor protein binding family B member 1

3

1

High-throughput

0.604

High

1395440

UBP16
Ubiquitin carboxyl-terminal hydrolase 16

3

4

High-throughput

0.604

High

1251592

HSDL2
Hydroxysteroid dehydrogenase-like protein 2

2

0

High-throughput

0.602

High

1395377

PRIPO
DNA-directed primase/polymerase protein

2

0

High-throughput

0.602

High

1048645

SSBP
Single-stranded DNA-binding protein, mitochondrial

2

0

High-throughput

0.585

High

914718

RM12
Large ribosomal subunit protein bL12m

2

0

High-throughput

0.576

High

1395384

MA7D2
MAP7 domain-containing protein 2

2

2

High-throughput

0.574

High

1210953

CE170
Centrosomal protein of 170 kDa

2

1

Small-scale

0.571

High

1215491

CE162
Centrosomal protein of 162 kDa

2

3

High-throughput

0.571

High

1350413

BICD2
Protein bicaudal D homolog 2

2

2

High-throughput

0.558

High

1395434

UBP20
Ubiquitin carboxyl-terminal hydrolase 20

2

4

Small-scale

0.558

High

322983

UBP19
Ubiquitin carboxyl-terminal hydrolase 19

2

2

High-throughput

0.549

High

698343

MAP4
Microtubule-associated protein 4

2

1

High-throughput

0.549

High

1309537

MAPK5
MAP kinase-activated protein kinase 5

2

1

High-throughput

0.549

High

414414

P53
Cellular tumor antigen p53

2

0

Small-scale

0.547

High

667409

RPB1
DNA-directed RNA polymerase II subunit RPB1

2

0

Small-scale

0.547

High

1186296

MA7D1
MAP7 domain-containing protein 1

2

2

Small-scale

0.521

High

1288757

KTN1
Kinectin

2

4

Small-scale

0.521

High

1239913

CAVN1
Caveolae-associated protein 1

1

0

High-throughput

0.513

High

1281314

RM10
Large ribosomal subunit protein uL10m

1

0

High-throughput

0.513

High

1347444

CEP76
Centrosomal protein of 76 kDa

1

0

Small-scale

0.513

High

1389256

RM53
Large ribosomal subunit protein mL53

1

0

High-throughput

0.513

High

1395401

MLP3B
Microtubule-associated protein 1 light chain 3 beta

1

1

High-throughput

0.501

High

49574

FEZF1
Fez family zinc finger protein 1

1

1

High-throughput

0.489

High

248755

PEPL
Periplakin

1

3

High-throughput

0.489

High

777541

K22E
Keratin, type II cytoskeletal 2 epidermal

1

2

High-throughput

0.489

High

851392

PRS6B
26S proteasome regulatory subunit 6B

1

2

High-throughput

0.489

High

997466

TBB4B
Tubulin beta-4B chain

1

2

High-throughput

0.489

High

1124523

MAP7
Ensconsin

1

3

High-throughput

0.489

High

1395382

SPAG5
Sperm-associated antigen 5

1

2

High-throughput

0.489

High

1395424

PNMA2
Paraneoplastic antigen Ma2

1

1

High-throughput

0.489

High

63312

ODAD3
Outer dynein arm-docking complex subunit 3

1

2

High-throughput

0.479

High

287996

MYCB2
E3 ubiquitin-protein ligase MYCBP2

1

3

High-throughput

0.479

High

1203227

ODFP2
Outer dense fiber protein 2

1

2

High-throughput

0.479

High

1209523

LRRK2
Leucine-rich repeat serine/threonine-protein kinase 2

1

3

Small-scale

0.479

High

1226222

CP135
Centrosomal protein of 135 kDa

1

3

High-throughput

0.479

High

1395395

CP250
Centrosome-associated protein CEP250

1

6

High-throughput

0.479

High

1395427

HECD1
E3 ubiquitin-protein ligase HECTD1

1

1

High-throughput

0.479

High

1143473

PCM1
Pericentriolar material 1 protein

1

0

Small-scale

0.458

High

1367027

ESR2
Estrogen receptor beta

1

0

High-throughput

0.458

High

1382332

RN166
E3 ubiquitin-protein ligase RNF166

1

0

High-throughput

0.458

High

1395407

MLP3A
Microtubule-associated protein 1 light chain 3 alpha

1

1

High-throughput

0.457

High

292198

S2512
Electrogenic aspartate/glutamate antiporter SLC25A12, mitochondrial

1

1

High-throughput

0.445

High

530228

TAU
Microtubule-associated protein tau

1

1

High-throughput

0.445

High

552058

ADT1
ADP/ATP translocase 1

1

1

High-throughput

0.445

High

749985

KINH
Kinesin-1 heavy chain

1

1

High-throughput

0.445

High

1217492

CE350
Centrosome-associated protein 350

1

3

High-throughput

0.445

High

1395392

TCHP
Trichoplein keratin filament-binding protein

1

2

High-throughput

0.445

High

1395399

UBE2O
(E3-independent) E2 ubiquitin-conjugating enzyme

1

4

High-throughput

0.445

High

1395414

SYIM
Isoleucine--tRNA ligase, mitochondrial

1

1

High-throughput

0.445

High

1395416

ATD3A
ATPase family AAA domain-containing protein 3A

1

1

High-throughput

0.445

High

1395425

MTUS1
Microtubule-associated tumor suppressor 1

1

3

High-throughput

0.445

High

1395437

UBP15
Ubiquitin carboxyl-terminal hydrolase 15

1

2

High-throughput

0.445

High

331241

KI20A
Kinesin-like protein KIF20A

1

3

High-throughput

0.437

High

350112

SNAI1
Zinc finger protein SNAI1

1

1

High-throughput

0.437

High

631413

RAB5A
Ras-related protein Rab-5A

1

1

Small-scale

0.437

High

956602

RAB5B
Ras-related protein Rab-5B

1

1

Small-scale

0.437

High

968784

PRS4
26S proteasome regulatory subunit 4

1

2

High-throughput

0.437

High

1351942

RBCC1
RB1-inducible coiled-coil protein 1

1

4

High-throughput

0.437

High

1353406

UBP33
Ubiquitin carboxyl-terminal hydrolase 33

1

3

Small-scale

0.437

High

1395403

CCDC8
Coiled-coil domain-containing protein 8

1

1

High-throughput

0.437

High

1395408

PKHA4
Pleckstrin homology domain-containing family A member 4

1

2

High-throughput

0.437

High

1395431

SMC3
Structural maintenance of chromosomes protein 3

1

4

High-throughput

0.437

High

359712

DHE3
Glutamate dehydrogenase 1, mitochondrial

1

0

High-throughput

0.426

High

457122

CAN1
Calpain-1 catalytic subunit

1

0

High-throughput

0.426

High

723053

RPB2
DNA-directed RNA polymerase II subunit RPB2

1

0

High-throughput

0.426

High

895282

CPT1A
Carnitine O-palmitoyltransferase 1, liver isoform

1

0

High-throughput

0.426

High

904363

BRCA2
Breast cancer type 2 susceptibility protein

1

0

High-throughput

0.426

High

1013434

LACTB
Serine beta-lactamase-like protein LACTB, mitochondrial

1

0

High-throughput

0.426

High

1058213

C1QBP
Complement component 1 Q subcomponent-binding protein, mitochondrial

1

0

High-throughput

0.426

High

1198497

CEP55
Centrosomal protein of 55 kDa

1

0

High-throughput

0.426

High

1326213

PNKD
Probable thioesterase PNKD

1

0

High-throughput

0.426

High

1395383

CK5P1
Mitochondrial tRNA methylthiotransferase CDK5RAP1

1

0

High-throughput

0.426

High

1395396

CA043
Protein C1orf43

1

0

High-throughput

0.426

High

1395410

PATZ1
POZ-, AT hook-, and zinc finger-containing protein 1

1

0

High-throughput

0.426

High

244180

KIF5C
Kinesin heavy chain isoform 5C

1

1

High-throughput

0.407

High

273573

DJC13
DnaJ homolog subfamily C member 13

1

1

High-throughput

0.407

High

300598

KRT36
Keratin, type I cuticular Ha6

1

3

High-throughput

0.407

High

334077

SMC2
Structural maintenance of chromosomes protein 2

1

3

High-throughput

0.407

High

407331

K2C6B
Keratin, type II cytoskeletal 6B

1

2

High-throughput

0.407

High

408136

TBB4A
Tubulin beta-4A chain

1

2

High-throughput

0.407

High

771805

SCN4A
Sodium channel protein type 4 subunit alpha

1

2

High-throughput

0.407

High

883934

CENPF
Centromere protein F

1

6

High-throughput

0.407

High

1093161

STK4
Serine/threonine-protein kinase 4

1

1

High-throughput

0.407

High

1106226

MYO9B
Unconventional myosin-IXb

1

4

High-throughput

0.407

High

1110536

KCC2D
Calcium/calmodulin-dependent protein kinase type II subunit delta

1

1

High-throughput

0.407

High

1127672

KT33B
Keratin, type I cuticular Ha3-II

1

4

High-throughput

0.407

High

1155138

SCN9A
Sodium channel protein type 9 subunit alpha

1

2

High-throughput

0.407

High

1156576

CCDC6
Coiled-coil domain-containing protein 6

1

2

High-throughput

0.407

High

1233135

RGPA1
Ral GTPase-activating protein subunit alpha-1

1

1

High-throughput

0.407

High

1293034

RLGPB
Ral GTPase-activating protein subunit beta

1

1

High-throughput

0.407

High

1336788

SMC1B
Structural maintenance of chromosomes protein 1B

1

3

High-throughput

0.407

High

1350551

NEK9
Serine/threonine-protein kinase Nek9

1

1

High-throughput

0.407

High

1395376

ASPP1
Apoptosis-stimulating of p53 protein 1

1

1

High-throughput

0.407

High

1395380

ALS2
Alsin

1

2

High-throughput

0.407

High

1395391

TBA1C
Tubulin alpha-1C chain

1

1

High-throughput

0.407

High

1395393

TBB6
Tubulin beta-6 chain

1

2

High-throughput

0.407

High

1395404

SLK
STE20-like serine/threonine-protein kinase

1

3

High-throughput

0.407

High

1395412

BIRC6
Dual E2 ubiquitin-conjugating enzyme/E3 ubiquitin-protein ligase BIRC6

1

3

High-throughput

0.407

High

1395413

KRT84
Keratin, type II cuticular Hb4

1

2

High-throughput

0.407

High

1395418

SCN3A
Sodium channel protein type 3 subunit alpha

1

2

High-throughput

0.407

High

1395419

SYLC
Leucine--tRNA ligase, cytoplasmic

1

1

High-throughput

0.407

High

1395428

MYO5B
Unconventional myosin-Vb

1

3

High-throughput

0.407

High

1395433

ZBTB1
Zinc finger and BTB domain-containing protein 1

1

1

High-throughput

0.407

High

1395438

MYO5A
Unconventional myosin-Va

1

3

High-throughput

0.407

High

1161185

MK06
Mitogen-activated protein kinase 6

7

0

Small-scale

0.392

High

1395421

DBNL
Drebrin-like protein

6

0

High-throughput

0.379

High

1395386

DNJC7
DnaJ homolog subfamily C member 7

4

0

High-throughput

0.360

High

239173

ZNRD2
Protein ZNRD2

4

0

Small-scale

0.353

High

1395397

SPZ1
Spermatogenic leucine zipper protein 1

3

0

High-throughput

0.347

High

1335301

FA98A
Protein FAM98A

3

0

High-throughput

0.345

High

1395400

PDGFD
Platelet-derived growth factor D

3

0

High-throughput

0.345

High

1348939

ZCH10
Zinc finger CCHC domain-containing protein 10

3

0

High-throughput

0.335

High

1033412

CDR2
Cerebellar degeneration-related protein 2

2

0

High-throughput

0.328

High

1063837

LG3BP
Galectin-3-binding protein

2

0

High-throughput

0.328

High

1312906

DRC2
Dynein regulatory complex subunit 2

2

0

High-throughput

0.328

High

1385012

ZN669
Zinc finger protein 669

2

0

High-throughput

0.328

High

1395411

Z286A
Zinc finger protein 286A

2

0

High-throughput

0.328

High

1395423

FBXL5
F-box/LRR-repeat protein 5

2

0

High-throughput

0.328

High

982774

RS24
Small ribosomal subunit protein eS24

2

0

High-throughput

0.313

High

1204387

SGO1
Shugoshin 1

2

0

High-throughput

0.313

High

1326033

RYBP
RING1 and YY1-binding protein

2

0

High-throughput

0.313

High

1395385

EIF3C
Eukaryotic translation initiation factor 3 subunit C

2

0

High-throughput

0.313

High

1395402

NAT10
RNA cytidine acetyltransferase

2

0

High-throughput

0.313

High

1395436

ZN451
E3 SUMO-protein ligase ZNF451

2

0

High-throughput

0.313

High

1065998

VAC14
Protein VAC14 homolog

2

0

Small-scale

0.298

High

96975

NPIB6
Nuclear pore complex-interacting protein family member B6

1

0

High-throughput

0.279

Low

155209

NCKP5
Nck-associated protein 5

1

0

High-throughput

0.279

Low

320586

FARP2
FERM, ARHGEF and pleckstrin domain-containing protein 2

1

0

High-throughput

0.279

Low

377291

GLUC
Pro-glucagon [Cleaved into: Glicentin; Glicentin-related polypeptide

1

0

High-throughput

0.279

Low

654128

XPA
DNA repair protein complementing XP-A cells

1

0

High-throughput

0.279

Low

848701

MAGA9
Melanoma-associated antigen 9

1

0

High-throughput

0.279

Low

1088786

NMDE1
Glutamate receptor ionotropic, NMDA 2A

1

0

High-throughput

0.279

Low

1098096

PSMD2
26S proteasome non-ATPase regulatory subunit 2

1

0

High-throughput

0.279

Low

1115914

DYR1A
Dual specificity tyrosine-phosphorylation-regulated kinase 1A

1

0

High-throughput

0.279

Low

1117254

NCOA4
Nuclear receptor coactivator 4

1

0

High-throughput

0.279

Low

1329409

SAMD3
Sterile alpha motif domain-containing protein 3

1

0

High-throughput

0.279

Low

1395389

RETNB
Resistin-like beta

1

0

High-throughput

0.279

Low

1395390

SOST
Sclerostin

1

0

High-throughput

0.279

Low

1395398

NUF2
Kinetochore protein Nuf2

1

0

High-throughput

0.279

Low

1395406

TXNIP
Thioredoxin-interacting protein

1

0

High-throughput

0.279

Low

1395409

CLXN
Calaxin

1

0

High-throughput

0.279

Low

1163972

DDR2
Discoidin domain-containing receptor 2

1

0

High-throughput

0.261

Low

67010

CO6A6

1

0

High-throughput

0.249

Low

260308

BRD4
Bromodomain-containing protein 4

1

0

High-throughput

0.249

Low

446458

G6PI
Glucose-6-phosphate isomerase

1

0

Small-scale

0.249

Low

504754

FBSP1
F-box/SPRY domain-containing protein 1

1

0

High-throughput

0.249

Low

584640

AREG
Amphiregulin

1

0

High-throughput

0.249

Low

589426

RFA2
Replication protein A 32 kDa subunit

1

0

Small-scale

0.249

Low

696229

RFA1
Replication protein A 70 kDa DNA-binding subunit

1

0

Small-scale

0.249

Low

768908

RFA3
Replication protein A 14 kDa subunit

1

0

Small-scale

0.249

Low

769763

RFC4
Replication factor C subunit 4

1

0

High-throughput

0.249

Low

932414

GALC
Galactocerebrosidase

1

0

High-throughput

0.249

Low

1250187

LRC31
Leucine-rich repeat-containing protein 31

1

0

High-throughput

0.249

Low

1395394

DUS26
Dual specificity protein phosphatase 26

1

0

High-throughput

0.249

Low

1395420

ZRAN1
Ubiquitin thioesterase ZRANB1

1

0

High-throughput

0.249

Low

66650

CC85C
Coiled-coil domain-containing protein 85C

1

0

High-throughput

0.232

Low

137087

PLOD2
Procollagen-lysine,2-oxoglutarate 5-dioxygenase 2

1

0

High-throughput

0.232

Low

192844

RPB4
DNA-directed RNA polymerase II subunit RPB4

1

0

High-throughput

0.232

Low

269881

IGSF3
Immunoglobulin superfamily member 3

1

0

High-throughput

0.232

Low

339566

ACSL3
Fatty acid CoA ligase Acsl3

1

0

High-throughput

0.232

Low

506021

GRL1A
DNA-directed RNA polymerase II subunit GRINL1A

1

0

High-throughput

0.232

Low

621715

RPB3
DNA-directed RNA polymerase II subunit RPB3

1

0

High-throughput

0.232

Low

622845

ERCC3
General transcription and DNA repair factor IIH helicase/translocase subunit XPB

1

0

High-throughput

0.232

Low

657920

SYWC
Tryptophan--tRNA ligase, cytoplasmic

1

0

High-throughput

0.232

Low

704985

RBL1
Retinoblastoma-like protein 1

1

0

High-throughput

0.232

Low

744683

TF2H1
General transcription factor IIH subunit 1

1

0

High-throughput

0.232

Low

787773

RPB9
DNA-directed RNA polymerase II subunit RPB9

1

0

High-throughput

0.232

Low

833672

NAA10
N-alpha-acetyltransferase 10

1

0

High-throughput

0.232

Low

839493

PI4KA
Phosphatidylinositol 4-kinase alpha

1

0

High-throughput

0.232

Low

854715

RAGP1
Ran GTPase-activating protein 1

1

0

High-throughput

0.232

Low

861252

MAP1B
Microtubule-associated protein 1B

1

0

High-throughput

0.232

Low

897066

MAP2
Methionine aminopeptidase 2

1

0

High-throughput

0.232

Low

903234

SMCA2
SWI/SNF-related matrix-associated actin-dependent regulator of chromatin subfamily A member 2

1

0

High-throughput

0.232

Low

912104

RPAB3
DNA-directed RNA polymerases I, II, and III subunit RPABC3

1

0

High-throughput

0.232

Low

912217

RPB11
DNA-directed RNA polymerase II subunit RPB11-a

1

0

High-throughput

0.232

Low

923523

RPAB4
DNA-directed RNA polymerases I, II, and III subunit RPABC4

1

0

High-throughput

0.232

Low

935009

TERA
Transitional endoplasmic reticulum ATPase

1

0

High-throughput

0.232

Low

976132

RPB7
DNA-directed RNA polymerase II subunit RPB7

1

0

High-throughput

0.232

Low

1035532

CENPE
Centromere-associated protein E

1

0

High-throughput

0.232

Low

1060563

KLC1
Kinesin light chain 1

1

0

High-throughput

0.232

Low

1089342

AIMP1
Aminoacyl tRNA synthase complex-interacting multifunctional protein 1

1

0

High-throughput

0.232

Low

1090173

PTN13
Tyrosine-protein phosphatase non-receptor type 13

1

0

High-throughput

0.232

Low

1095218

MTAP
S-methyl-5'-thioadenosine phosphorylase

1

0

High-throughput

0.232

Low

1096617

AIMP2
Aminoacyl tRNA synthase complex-interacting multifunctional protein 2

1

0

High-throughput

0.232

Low

1149456

SKI2
Superkiller complex protein 2

1

0

High-throughput

0.232

Low

1162656

DGKD
Diacylglycerol kinase delta

1

0

High-throughput

0.232

Low

1178684

RHG31
Rho GTPase-activating protein 31

1

0

High-throughput

0.232

Low

1193661

MAP9
Microtubule-associated protein 9

1

0

High-throughput

0.232

Low

1226794

AR6P4
ADP-ribosylation factor-like protein 6-interacting protein 4

1

0

High-throughput

0.232

Low

1244368

SKI3
Superkiller complex protein 3

1

0

High-throughput

0.232

Low

1277703

MYCPP
C-myc promoter-binding protein

1

0

High-throughput

0.232

Low

1287203

PEG10
Retrotransposon-derived protein PEG10

1

0

High-throughput

0.232

Low

1311375

ANKH1
Ankyrin repeat and KH domain-containing protein 1

1

0

High-throughput

0.232

Low

1313033

RPAP2
Putative RNA polymerase II subunit B1 CTD phosphatase RPAP2

1

0

High-throughput

0.232

Low

1323803

ARG28
Rho guanine nucleotide exchange factor 28

1

0

High-throughput

0.232

Low

1327647

NKAP
NF-kappa-B-activating protein

1

0

High-throughput

0.232

Low

1337151

ABCF1
ATP-binding cassette sub-family F member 1

1

0

High-throughput

0.232

Low

1338754

ARI1B
AT-rich interactive domain-containing protein 1B

1

0

High-throughput

0.232

Low

1367959

TF2H4
General transcription factor IIH subunit 4

1

0

High-throughput

0.232

Low

1369129

COR2A
Coronin-2A

1

0

High-throughput

0.232

Low

1370739

SMRD2
SWI/SNF-related matrix-associated actin-dependent regulator of chromatin subfamily D member 2

1

0

High-throughput

0.232

Low

1371502

USP9X
Ubiquitin carboxyl-terminal hydrolase 9X

1

0

High-throughput

0.232

Low

1389136

F120B
Constitutive coactivator of peroxisome proliferator-activated receptor gamma

1

0

High-throughput

0.232

Low

1395378

KLH15
Kelch-like protein 15

1

0

High-throughput

0.232

Low

1395379

MF14A
Hippocampus abundant transcript 1 protein

1

0

High-throughput

0.232

Low

1395381

TRPM7
Transient receptor potential cation channel subfamily M member 7

1

0

High-throughput

0.232

Low

1395387

CSN8
COP9 signalosome complex subunit 8

1

0

High-throughput

0.232

Low

1395388

RGPD5
RANBP2-like and GRIP domain-containing protein 5/6

1

0

High-throughput

0.232

Low

1395405

ZN106
Zinc finger protein 106

1

0

High-throughput

0.232

Low

1395415

SEP11
Septin-11

1

0

High-throughput

0.232

Low

1395417

GFOD1
Glucose-fructose oxidoreductase domain-containing protein 1

1

0

High-throughput

0.232

Low

1395422

FBX3
F-box only protein 3

1

0

High-throughput

0.232

Low

1395426

MED23
Mediator of RNA polymerase II transcription subunit 23

1

0

High-throughput

0.232

Low

1395429

SCML1
Sex comb on midleg-like protein 1

1

0

High-throughput

0.232

Low

1395430

CSN3
COP9 signalosome complex subunit 3

1

0

High-throughput

0.232

Low

1395432

FYV1
1-phosphatidylinositol 3-phosphate 5-kinase

1

0

High-throughput

0.232

Low

1395435

S27A6
Long-chain fatty acid transport protein 6

1

0

High-throughput

0.232

Low

1395439

NDK7
Nucleoside diphosphate kinase homolog 7

1

0

High-throughput

0.232

Low


Service provided by Combinatics, Japan

Last updated: 11Aug2025